When initially described in 2009, Titanoboa was estimated to have been about 12.8 meters long. This meant that in at least terms of length, Titanoboa was larger than the previous record holder for largest ever snake, Gigantophis, by a comfortable margin. Later modelling suggested a total length of about 14.6 meters, a figure that has since been commonly rounded off to 15 meters by others.
Reptiles are commonly thought to grow in accordance with the available ambient temperature of a climate. This is because higher temperatures that remain fairly constant throughout the year with very little seasonal variation allow ectothermic (cold blooded) animals to maintain an optimum metabolism for longer. This means that bodily functions such as digestion, circulation and respiration among others all become far more efficient, and a greater amount of energy can be set aside for other areas such as growth.
In reference to the larger size of Titanoboa, this could indicate that sixty million years ago global temperatures, specifically at the equator but quite possibly further away, were considerably higher than those we know today. This is because today the largest known snakes which live close to the equator can commonly attain sizes of five to six meters in length, with rarer individuals approaching six and a half to seven meters long.
Titanoboa fossils are so far only known from the Cerrejón Formation of Colombia in South America. The Cerrejón Formation represents what is at the time of writing the earliest known occurrence of Neotropical rainforests (rainforests of Central and South America). The area of the Cerrejón Formation that the Titanoboa holotype fossils are known from has been established as going back to the Selandian of the Paleocene. This means that Titanoboa are known to have lived about sixty million years ago (give or a take a million years), and approximately five million years after the KT extinction which marks the end of the Mesozoic and the disappearance of the dinosaurs. During this time, Titanoboa would have lived and hunted in low lying rainforests that contained an extensive system of rivers that criss-crossed over the landscape.
Titanoboa would have been at their most dangerous when in the water. When in the water, body weight means very little since the buoyancy of the water would counteract the effects of gravity upon the body (this is exactly why marine animals like whales grow so big). This would mean that even a large Titanoboa would have been very quick when moving through the water as well as expending comparatively little energy to do so than what it would have had to do if on land. Another advantage of hunting in the water is that the sheer bulk of the body of Titanoboa would have been hidden by the water. When striking animals that were on or near the surface, the surface sheen of the water would have hidden any approach from a Titanoboa submerged under the surface, while the Titanoboa would have been able to lock on to the silhouette of its targeted prey. In addition Titanoboa would have been capable of lurking upon the bottom of water system and holding its breath for a considerable time, waiting for other animals swimming through the water which it could then strike at from below.
And to please @Slogokok34 I make a small change to Gen 2 by including a size reference chart.
Metahub Tier: TBA.
Critical chance: 20%
Shielded Decelerating Strike.
Immune to Distraction.
On Escape Dust Cloud.